{# load django_bootstrap5#%} CCDB server
Prof. Itay Mayrose Lab - Plant Evolution, bioinformatics, & comparative genomics


   CCDB is a community resource. If you are aware of any missing data or if any of the counts are erroneous please submit it to the forms submit new counts, or report erroneous counts.


CCDBAngiospermsBoraginaceaeSymphytumSymphytum officinale

123 chromosome counts in Symphytum officinale:


Name Accepted Name Gametophytic(n) Sporophytic(2n) Data Source reference
!   Symphytum officinale Symphytum officinale L.   36 Darlington, 1955 Suzuka 1950
  Symphytum officinale L. Symphytum officinale L.   24 CHROBASE GADELLA T.W.J., KLIPHUIS E. - 1970. Cytotaxonomic investigations in some Angiosperms collected in the valley of Aosta and in the National Park Gran Paradiso.. Caryologia , 23(3): 363-379.
  Symphytum officinale L. Symphytum officinale L.   24 CHROBASE Bottega S., Garbari F., Peruzzi L. - 2000. Aspetti biosistematici di popolazioni toscane afferenti a Symphytum gr. officinale (Boraginaceae). "Problematiche di Biologia Vegetale in Ambiente Mediterraneo", 95 Congresso della S.B.I. , : 71.
  Symphytum officinale L. Symphytum officinale L.   24 CHROBASE Bottega S., Garbari F., Peruzzi L. - 2000. Aspetti biosistematici di popolazioni toscane afferenti a Symphytum gr. officinale (Boraginaceae). "Problematiche di Biologia Vegetale in Ambiente Mediterraneo", 95 Congresso della S.B.I. , : 71.
  Symphytum officinale L. Symphytum officinale L.   48 CHROBASE PERUZZI L., GARBARI F., BOTTEGA S. - 2001. Symphytum tanaicense Steven (Boraginaceae) new for the Italian Flora. Willdenowia , 31(1): 33-41.
  Symphytum officinale L. Symphytum officinale L.   24 CHROBASE PERUZZI L., GARBARI F., BOTTEGA S. - 2001. Symphytum tanaicense Steven (Boraginaceae) new for the Italian Flora. Willdenowia , 31(1): 33-41.
  Symphytum officinale L. Symphytum officinale L.   24 CHROBASE BOTTEGA S., GARBARI F., PERUZZI L. - 2001. Mediterranean chromosome number reports - 11.. Flora Mediterranea , 11: 436-439.
  Symphytum officinale L. Symphytum officinale L.   36 Slovakia Database Májovský J. et al., 1974a: Index of chromosome numbers of Slovakian flora (Part 3). - Acta Fac. Rerum Nat. Univ. Comen., Bot. 22: 1-20.
  Symphytum officinale L. Symphytum officinale L.   48 Slovakia Database Migra V., 1982: Zytotaxonomische Probleme ausgewählter Taxa der Flora des Babia Hora - Bergmassivs (Gebirge Západné Beskydy) I. - Acta Fac. Rerum Nat. Univ. Comen., Bot. 29: 87-95.
  Symphytum officinale L. Symphytum officinale L.   48 Slovakia Database Murín A. & Májovský J., 1982: Die Bedeutung der Polyploidie in der Entwicklung der in der Slowakei wachsenden Arten der Gattung Symphytum L. - Acta Fac. Rerum Nat. Univ. Comen., Bot. 29: 1-25.
  Symphytum officinale L. Symphytum officinale L.   48 Slovakia Database Murín A. & Májovský J., 1982: Die Bedeutung der Polyploidie in der Entwicklung der in der Slowakei wachsenden Arten der Gattung Symphytum L. - Acta Fac. Rerum Nat. Univ. Comen., Bot. 29: 1-25.
  Symphytum officinale L. Symphytum officinale L.   48 Slovakia Database Murín A. & Májovský J., 1982: Die Bedeutung der Polyploidie in der Entwicklung der in der Slowakei wachsenden Arten der Gattung Symphytum L. - Acta Fac. Rerum Nat. Univ. Comen., Bot. 29: 1-25.
  Symphytum officinale L. Symphytum officinale L.   48 Slovakia Database Murín A. & Májovský J., 1982: Die Bedeutung der Polyploidie in der Entwicklung der in der Slowakei wachsenden Arten der Gattung Symphytum L. - Acta Fac. Rerum Nat. Univ. Comen., Bot. 29: 1-25.
  Symphytum officinale L. Symphytum officinale L.   48 Slovakia Database Murín A. & Májovský J., 1982: Die Bedeutung der Polyploidie in der Entwicklung der in der Slowakei wachsenden Arten der Gattung Symphytum L. - Acta Fac. Rerum Nat. Univ. Comen., Bot. 29: 1-25.
  Symphytum officinale L. Symphytum officinale L.   48 Slovakia Database Murín A. & Májovský J., 1982: Die Bedeutung der Polyploidie in der Entwicklung der in der Slowakei wachsenden Arten der Gattung Symphytum L. - Acta Fac. Rerum Nat. Univ. Comen., Bot. 29: 1-25.
  Symphytum officinale L. Symphytum officinale L.   48 Slovakia Database Murín A. & Májovský J., 1982: Die Bedeutung der Polyploidie in der Entwicklung der in der Slowakei wachsenden Arten der Gattung Symphytum L. - Acta Fac. Rerum Nat. Univ. Comen., Bot. 29: 1-25.
!   Symphytum officinale L. Symphytum officinale L.   24 British Isles cytology Gadella, T.W.J., Kliphuis, E., Perring, F.H. Acta Bot. Neerl. 23: 433-437 (1974)
!   Symphytum officinale L. Symphytum officinale L.   44 British Isles cytology Gadella, T.W.J., Kliphuis, E., Perring, F.H. Acta Bot. Neerl. 23: 433-437 (1974)
!   Symphytum officinale L. Symphytum officinale L.   48 British Isles cytology Gadella, T.W.J., Kliphuis, E., Perring, F.H. Acta Bot. Neerl. 23: 433-437 (1974)
!   Symphytum officinale L. Symphytum officinale L.   48 British Isles cytology Marks, G.E. (1975)
!   Symphytum officinale L. Symphytum officinale L.   44 rare, 48 flora of NW Europe http://wbd.etibioinformatics.nl/bis/flora.php?selected=beschrijving&menuentry=soorten&id=3415
!   Symphytum officinale L. Symphytum officinale L.   54 Flora Europaea Markova & Ivanova,
!   Symphytum officinale L. Symphytum officinale L.   24 + 4B Flora Europaea Gadella & Kliphuis
!   Symphytum officinale L. Symphytum officinale L.   24 Poland Database Skali?ska M., Jankun A., Wcis?o H. et al. 1971. Studies in chromosome numbers of Polish Angiosperms. Eighth contribution. Acta Biol. Cracov. Ser. Bot. 14, 55-102.
!   Symphytum officinale L. Symphytum officinale L.   48 Poland Database Skali?ska M., Jankun A., Wcis?o H. et al. 1971. Studies in chromosome numbers of Polish Angiosperms. Eighth contribution. Acta Biol. Cracov. Ser. Bot. 14, 55-102.
  Symphytum officinale L. Symphytum officinale L.   48 IPCN online Dmitrieva, S. A. & V. I. Parfenov. 1985. Kariologicheskaja kharakteristika nekotorykh vidov poleznykh rastenij flory Belorussii. Izv. Akad. Nauk Belorussk. SSR, Ser. Biol. Nauk 6: 3–8.
  Symphytum officinale L. Symphytum officinale L.   40-48 IPCN online Gadella, T. W. & E. KLIPHUIS. 1984. Population variability, hybridization and introgression in Symphytum officinale L. in the Netherlands. Bot. Jahrb. Syst. 104: 519–536.
  Symphytum officinale L. Symphytum officinale L.   24,48 IPCN online Murin, A. & J. Majovsky. 1982. Die Bedeutung der Polyploidie in der Entwicklung in der Slowakei wachsenden Arten der Gattung Symphytum L. Acta Fac. Rerum Nat. Univ. Comenianae, Bot. 29: 1–25.
  Symphytum officinale L. Symphytum officinale L.   45 IPCN online SHIRATO, K., T. Shintani, G. Nakanishi & A. Kamizy;afo. 1985. Chromosome numbers in genus Symphytum. Chromosome Inf. Serv. 38: 21–23.
  Symphytum officinale L. Symphytum officinale L.   48 IPCN online Sekovski, Z. & M. Jovanovska. 1983. Chromosome atlas of some Macedonian angiosperms. IV. Ann. Fac. Biol. Univ. Skopje. 36: 73–86.
  Symphytum officinale L. Symphytum officinale L.   24, 48 IPCN online Gadella, T. W. & E. KLIPHUIS. 1985. Diploide populaties van de Smeerwortel in Nederland bedreigd. Gorteria 12: 156–158.
  Symphytum officinale L. Symphytum officinale L.   24, 40, 48, 56 IPCN online Huizing, H. J. 1985. Phytochemistry, systematics and biogenesis of pyrrolizidine alkaloids of Symphytum taxa. Thesis, Meppel. ý.
  Symphytum officinale L. Symphytum officinale L.   48 IPCN online Gadella, T. W. 1983. Enkele floristische en cytotaxonomische waaremingen op Schiermonnikoog. Gorteria 10: 227–229.
  Symphytum officinale L. Symphytum officinale L.   24,40,48 IPCN online Huizing, H. J., T. W. J. Gadella & E. Kliphuis. 1982. Chemotaxonomical investigations of the Symphytum officinale polyploid complex and S. asperum (Boraginaceae): The pyrrolizidine alkaloids. Pl. Syst. Evol. 140: 279–292.
  Symphytum officinale L. Symphytum officinale L.   48 IPCN online Krahulcová, A. 1988. Selected chromosome counts of the Czechoslovak flora I. Folia Geobot. Phytotax. 23: 375–381.
  Symphytum officinale L. Symphytum officinale L.   24, 48 IPCN online Jaarsma, T. A., J. C. v. Loon, H. J. Huizing, T. M. Malingré & T. W. J. Gadella. 1989. Somaclonal variation in regenerated plants of Symphytum officinale L. Proc. Kon. Ned. Akad. Wetensch., C 92: 179–187.
  Symphytum officinale L. Symphytum officinale L.   24 IPCN online Mekki, L., H. t. Hart, N. Z. El-Alfy, A. Dewedar & T. W. J. Gadella. 1987. The Giemsa C-banded karyotype of diploid Symphytum officinale (Boraginaceae). Acta Bot. Neerl. 36: 33–37.
  Symphytum officinale L. Symphytum officinale L.   48 IPCN online Parfenov, V. I. & S. A. Dmitrieva. 1988. Kariologicheskaja kharakteristika predstavitelej flory sosudistykh rastenij Berezinskogo biosfernogo zapovednika. Zapov. Belorussii Issl. 12: 3–8.
  Symphytum officinale L. Symphytum officinale L.   24,40,48,56 IPCN online Van Loon, J. C. & B. Kieft. 1980. In Chromosome number reports LXVIII. Taxon 29: 538–542.
  Symphytum officinale L. Symphytum officinale L.   24,48 IPCN online PERRING, F. H. 1975. Network research, Symphytum survey. Watsonia 10: 296–297.
  Symphytum officinale L. Symphytum officinale L.   24,48,40,56 IPCN online Gadella, T. W. J. & E. Kliphuis. 1978. Cytotaxonomic studies in the genus Symphyrum. VIII. Chromosome numbers and classification of ten European species. Proc. Kon. Ned. Akad. Wetensch., C 81: 162–172.
  Symphytum officinale L. Symphytum officinale L.   40,48 IPCN online Gadella, T. W. J. & E. Kliphuis. 1978. Cytotaxonomic studies in the genus Symphyrum. VIII. Chromosome numbers and classification of ten European species. Proc. Kon. Ned. Akad. Wetensch., C 81: 162–172.
  Symphytum officinale L. Symphytum officinale L.   24+0-4B, 48 IPCN online Peruzzi, L., F. Garbari & S. Bottega. 2001. Symphytum tanaicense (Boraginaceae) new for the Italian flora. Willdenowia 31: 33–41.
  Symphytum officinale L. Symphytum officinale L.   24+0-4B IPCN online Bottega, S., F. Garbari & L. Peruzzi. 2001. Mediterranean chromosome number reports 11 (1227--1231). Fl. Medit. 11: 436–439.
  Symphytum officinale L. Symphytum officinale L.   48 IPCN online Dmitrieva, S. A. & V. I. Parfenov. 1985. Kariologicheskaja kharakteristika nekotorykh vidov poleznykh rastenij flory Belorussii. Izv. Akad. Nauk Belorussk. SSR, Ser. Biol. Nauk 6: 3–8.
  Symphytum officinale L. Symphytum officinale L.   40-48 IPCN online Gadella, T. W. & E. KLIPHUIS. 1984. Population variability, hybridization and introgression in Symphytum officinale L. in the Netherlands. Bot. Jahrb. Syst. 104: 519–536.
  Symphytum officinale L. Symphytum officinale L.   24,48 IPCN online Murin, A. & J. Majovsky. 1982. Die Bedeutung der Polyploidie in der Entwicklung in der Slowakei wachsenden Arten der Gattung Symphytum L. Acta Fac. Rerum Nat. Univ. Comenianae, Bot. 29: 1–25.
  Symphytum officinale L. Symphytum officinale L.   45 IPCN online SHIRATO, K., T. Shintani, G. Nakanishi & A. Kamizy;afo. 1985. Chromosome numbers in genus Symphytum. Chromosome Inf. Serv. 38: 21–23.
  Symphytum officinale L. Symphytum officinale L.   48 IPCN online Sekovski, Z. & M. Jovanovska. 1983. Chromosome atlas of some Macedonian angiosperms. IV. Ann. Fac. Biol. Univ. Skopje. 36: 73–86.
  Symphytum officinale L. Symphytum officinale L.   24, 48 IPCN online Gadella, T. W. & E. KLIPHUIS. 1985. Diploide populaties van de Smeerwortel in Nederland bedreigd. Gorteria 12: 156–158.
  Symphytum officinale L. Symphytum officinale L.   24, 40, 48, 56 IPCN online Huizing, H. J. 1985. Phytochemistry, systematics and biogenesis of pyrrolizidine alkaloids of Symphytum taxa. Thesis, Meppel. ý.
  Symphytum officinale L. Symphytum officinale L.   48 IPCN online Gadella, T. W. 1983. Enkele floristische en cytotaxonomische waaremingen op Schiermonnikoog. Gorteria 10: 227–229.
  Symphytum officinale L. Symphytum officinale L.   24,40,48 IPCN online Huizing, H. J., T. W. J. Gadella & E. Kliphuis. 1982. Chemotaxonomical investigations of the Symphytum officinale polyploid complex and S. asperum (Boraginaceae): The pyrrolizidine alkaloids. Pl. Syst. Evol. 140: 279–292.
  Symphytum officinale L. Symphytum officinale L.   48 IPCN online Krahulcová, A. 1988. Selected chromosome counts of the Czechoslovak flora I. Folia Geobot. Phytotax. 23: 375–381.
  Symphytum officinale L. Symphytum officinale L.   24, 48 IPCN online Jaarsma, T. A., J. C. v. Loon, H. J. Huizing, T. M. Malingré & T. W. J. Gadella. 1989. Somaclonal variation in regenerated plants of Symphytum officinale L. Proc. Kon. Ned. Akad. Wetensch., C 92: 179–187.
  Symphytum officinale L. Symphytum officinale L.   24 IPCN online Mekki, L., H. t. Hart, N. Z. El-Alfy, A. Dewedar & T. W. J. Gadella. 1987. The Giemsa C-banded karyotype of diploid Symphytum officinale (Boraginaceae). Acta Bot. Neerl. 36: 33–37.
  Symphytum officinale L. Symphytum officinale L.   48 IPCN online Parfenov, V. I. & S. A. Dmitrieva. 1988. Kariologicheskaja kharakteristika predstavitelej flory sosudistykh rastenij Berezinskogo biosfernogo zapovednika. Zapov. Belorussii Issl. 12: 3–8.
  Symphytum officinale L. Symphytum officinale L.   24,40,48,56 IPCN online Van Loon, J. C. & B. Kieft. 1980. In Chromosome number reports LXVIII. Taxon 29: 538–542.
  Symphytum officinale L. Symphytum officinale L.   24,48 IPCN online PERRING, F. H. 1975. Network research, Symphytum survey. Watsonia 10: 296–297.
  Symphytum officinale L. Symphytum officinale L.   24,48,40,56 IPCN online Gadella, T. W. J. & E. Kliphuis. 1978. Cytotaxonomic studies in the genus Symphyrum. VIII. Chromosome numbers and classification of ten European species. Proc. Kon. Ned. Akad. Wetensch., C 81: 162–172.
  Symphytum officinale L. Symphytum officinale L.   40,48 IPCN online Gadella, T. W. J. & E. Kliphuis. 1978. Cytotaxonomic studies in the genus Symphyrum. VIII. Chromosome numbers and classification of ten European species. Proc. Kon. Ned. Akad. Wetensch., C 81: 162–172.
  Symphytum officinale L. Symphytum officinale L.   24+0-4B, 48 IPCN online Peruzzi, L., F. Garbari & S. Bottega. 2001. Symphytum tanaicense (Boraginaceae) new for the Italian flora. Willdenowia 31: 33–41.
  Symphytum officinale L. Symphytum officinale L.   24+0-4B IPCN online Bottega, S., F. Garbari & L. Peruzzi. 2001. Mediterranean chromosome number reports 11 (1227--1231). Fl. Medit. 11: 436–439.
  Symphytum officinale L. Symphytum officinale L.   24, 48 IPCN72 BASLER, A. 1972. Cytotaxonomische Untersuchungen an der Boraginaceen-Gattung Symphytum L. Bot. Jahrb. 92: 508-553.
  Symphytum officinale L. Symphytum officinale L. 24   IPCN72 DELAY, J. 1972. Boraginacees (suite). Inf. Ann. Caryosyst. et Cytogenet. 6: 9-12.
  Symphytum officinale L. Symphytum officinale L.   24, 48 IPCN72 GADELLA, T.W.J. & E. KLIPHUIS. 1972. Cytotaxonomic studies in the genus Symphytum. IV. Cytogeographic investigations in Symphytum officinale L. Acta Bot. Neer!. 21: 169-173.
  Symphytum officinale L. Symphytum officinale L.   48 IPCN72 GADELLA, T., & E. KLIPHUIS. 1972. studies in chromosome numbers of Yugoslavian angiosperms. Acta Bot. Croatica 31: 91-103.
  Symphytum officinale L. Symphytum officinale L.   24, 48 IPCN72 WCISLO, H. 1972. Karyological studies in symphytum L. Acta Biol. Cracov. Ser. Bot. 15: 153-164.
  Symphytum officinale L. Symphytum officinale L.   24, 40, 48, 36, 44 IPCN67-71 GADELLA, T.W.J., & E. KLIPHUIS. 1969. Cytotaxonomic studies in the genus symphytum II. Crossing experiments between symphytum officinal L. and symphytum asperum Lepech. Acta Bot. Neerl. 18: 544-549.
  Symphytum officinale L. Symphytum officinale L. 18   IPCN67-71 LAANE, M.M. 1969. Meiosis and structural hybridity in some Norwegian plant species. Blyttia 27: 141-173.
  Symphytum officinale L. Symphytum officinale L.   24+4B IPCN67-71 GADELLA, TH. W.J., & E. KLIPHUIS. 1970. Cytotaxonomic investigations in some angiosperms collected in the Valley of Aosta and in the National Park "Gran Paradiso". Carylogia 23: 363-379.
  Symphytum officinale L. Symphytum officinale L.   48 IPCN67-71 GADELLA, TH. W.J., E. KLIPHUIS, & K.U. KRAMER. 1970. Zytotaxonomische Untersuchungen an Blutenpflanzen aus dem Osten Osterreichs. Wiss. Arbeiten Burgenland 44: 187-195.
  Symphytum officinale L. Symphytum officinale L. 12   IPCN67-71 LEPPER, L. 1970. Beitrage zur Chromosomenzahlen-Dokumentation. Wiss. z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 19: 369-376.
  Symphytum officinale L. Symphytum officinale L.   "24, 24+1-4B, 40-48*" IPCN67-71 GADELLA, Th. W. J., & E. KLIPHUIS. 1967. Chromosome numbers of flowering plants in the Netherlands III. Proc. Roy. Neth. Acad. Sci. Ser. c. 70: 7-20.
  Symphytum officinale L. Symphytum officinale L.   "24, 24+1-4B, 40-48*" IPCN67-71 GADELLA, Th. w. J . & E. KLIPHUIS. 1967. Cytotaxonomic studies in the genus Symphytum I. Symphytum officinale in the Netherlands. Proc. Roy. Neth. Acad. Sci. Ser. C. 70: 378 391.
  Symphytum officinale L. Symphytum officinale L.   40, 48 IPCN67-71 GADELLA, T.W.J., & E. KLIPHUIS. 1971. Cytotaxonomic studies in the genus Symphytum III. Some Symphytum hybrids in Belgium and the Netherlands. Biol. Jaarboek Dodonaea (Gent) 39: 97-107.
  Symphytum officinale L. Symphytum officinale L.   40 IPCN73-74 GADELLA, T.W.J., & E. KLIPHUIS. 1973. Cytotaxonomic studies in the genus symphytum. v. some notes on W. European plants with the chromosome number 2n=40. Bot. Jahrb. 93: 530-538. 1973.
  Symphytum officinale L. Symphytum officinale L.   40, 48 IPCN73-74 GADELLA, T.W.J., & E. KLIPHUIS. 1973. Chromosome numbers of flowerinl plants in the Netherlands VI. Proc. Kon. Ned. Acad. Wet., ser. C 76: 303-311.
  Symphytum officinale L. Symphytum officinale L.   24, 44, 48 IPCN73-74 GADELLA, T.W.J., E. KLIPHUIS, & F.H. PERRING. 1974. Cytotaxonomic studies in the genus Symphytum. VI. some notes on Symphytum in Britai Acta Bot. Neerl. 23: 433 437.
  Symphytum officinale L. Symphytum officinale L.   36 IPCN73-74 MAJOVSKY, J., ET AL. 1974. Index of chromosome numbers of Slovakian flora. (Part 3). Acta Fac. Rerum Nat. Univ. Comenianae Bot. 22: 1-20.
  Symphytum officinale L. Symphytum officinale L.   24, 48 IPCN73-74 PERRING, F.H. 1974. Symphytum survey. Watsonia 10: 296-297.
  Symphytum officinale L. Symphytum officinale L.   48 IPCN73-74 MARKOVA, M.L., & P. IVANOVA. 1974. In IOPB chromosome number report' XLIV. Taxon 23: 373-380.
!   Symphytum officinale L. Symphytum officinale L.   24 eFlora  
  Symphytum officinale L. Symphytum officinale L.   40, 48 Fedorov, 1974 Gadella, Kliphuis 19S3
  Symphytum officinale L. Symphytum officinale L.   40, 48 Fedorov, 1974 Tischler 1934
  Symphytum officinale L. Symphytum officinale L.   ~40 Fedorov, 1974 Liive A.;. Liive D. 1956b
  Symphytum officinale L. Symphytum officinale L.   48 Fedorov, 1974 Tamavschi 1948
  Symphytum officinale L. Symphytum officinale L.   48 Fedorov, 1974 Gadella, Kliphuis 1967
!   Symphytum officinale L. Symphytum officinale L.   2n = 48 Austria_checklist Gadella, T. W. J., Kliphuis, E., & Kramer, K. U. 1970. Zytotaxonomische Untersuchungen an Bl?tenpflanzen aus dem Osten ?sterreichs. Wiss.Arb.Burgenland 44, 187-195., Gadella, T. W. J. & Kliphuis, E. 1972. Cytotaxonomic studies in the genus Symphytum IV. Cytogeographic investigations in Symphytum officinale L. Acta Bot.Neerl. 21[2], 169-173., Jaarsma, T. A., Lohmanns, E., Gadella, T. W. J. et al. 1989. Chemotaxonomy of the Symphytum officinale agg. (Boraginaceae). Pl.Syst.Evol. 167[3/4], 113-127.
!   Symphytum officinale L. Symphytum officinale L.   2n = 48 Austria_checklist Gadella, T. W. J., Kliphuis, E., & Kramer, K. U. 1970. Zytotaxonomische Untersuchungen an Bl?tenpflanzen aus dem Osten ?sterreichs. Wiss.Arb.Burgenland 44, 187-195., Gadella, T. W. J. & Kliphuis, E. 1972. Cytotaxonomic studies in the genus Symphytum IV. Cytogeographic investigations in Symphytum officinale L. Acta Bot.Neerl. 21[2], 169-173., Jaarsma, T. A., Lohmanns, E., Gadella, T. W. J. et al. 1989. Chemotaxonomy of the Symphytum officinale agg. (Boraginaceae). Pl.Syst.Evol. 167[3/4], 113-127.
!   Symphytum officinale L. Symphytum officinale L.   2n = 48 Austria_checklist Gadella, T. W. J., Kliphuis, E., & Kramer, K. U. 1970. Zytotaxonomische Untersuchungen an Bl?tenpflanzen aus dem Osten ?sterreichs. Wiss.Arb.Burgenland 44, 187-195., Gadella, T. W. J. & Kliphuis, E. 1972. Cytotaxonomic studies in the genus Symphytum IV. Cytogeographic investigations in Symphytum officinale L. Acta Bot.Neerl. 21[2], 169-173.
!   Symphytum officinale L. Symphytum officinale L.   2n = 48 Austria_checklist Gadella, T. W. J., Kliphuis, E., & Kramer, K. U. 1970. Zytotaxonomische Untersuchungen an Bl?tenpflanzen aus dem Osten ?sterreichs. Wiss.Arb.Burgenland 44, 187-195., Gadella, T. W. J. & Kliphuis, E. 1972. Cytotaxonomic studies in the genus Symphytum IV. Cytogeographic investigations in Symphytum officinale L. Acta Bot.Neerl. 21[2], 169-173.
!   Symphytum officinale L. Symphytum officinale L.   2n = 48 Austria_checklist Gadella, T. W. J., Kliphuis, E., & Kramer, K. U. 1970. Zytotaxonomische Untersuchungen an Bl?tenpflanzen aus dem Osten ?sterreichs. Wiss.Arb.Burgenland 44, 187-195., Gadella, T. W. J. & Kliphuis, E. 1972. Cytotaxonomic studies in the genus Symphytum IV. Cytogeographic investigations in Symphytum officinale L. Acta Bot.Neerl. 21[2], 169-173.
!   Symphytum officinale L. Symphytum officinale L.   2n = 48 Austria_checklist Gadella, T. W. J., Kliphuis, E., & Kramer, K. U. 1970. Zytotaxonomische Untersuchungen an Bl?tenpflanzen aus dem Osten ?sterreichs. Wiss.Arb.Burgenland 44, 187-195., Gadella, T. W. J. & Kliphuis, E. 1972. Cytotaxonomic studies in the genus Symphytum IV. Cytogeographic investigations in Symphytum officinale L. Acta Bot.Neerl. 21[2], 169-173.
!   Symphytum officinale L. Symphytum officinale L.   2n = [48]U Austria_checklist Gadella, T. W. J. & Kliphuis, E. 1972. Cytotaxonomic studies in the genus Symphytum IV. Cytogeographic investigations in Symphytum officinale L. Acta Bot.Neerl. 21[2], 169-173.
!   Symphytum officinale L. Symphytum officinale L.   2n = [48]U Austria_checklist Gadella, T. W. J. & Kliphuis, E. 1972. Cytotaxonomic studies in the genus Symphytum IV. Cytogeographic investigations in Symphytum officinale L. Acta Bot.Neerl. 21[2], 169-173.
!   Symphytum officinale L. Symphytum officinale L.   2n = [48]U Austria_checklist Gadella, T. W. J. & Kliphuis, E. 1972. Cytotaxonomic studies in the genus Symphytum IV. Cytogeographic investigations in Symphytum officinale L. Acta Bot.Neerl. 21[2], 169-173.
!   Symphytum officinale L. Symphytum officinale L.   2n = [48]U Austria_checklist Gadella, T. W. J. & Kliphuis, E. 1972. Cytotaxonomic studies in the genus Symphytum IV. Cytogeographic investigations in Symphytum officinale L. Acta Bot.Neerl. 21[2], 169-173.
  Symphytum officinale L. Symphytum officinale L.   c.40 Cave1956 Love & Love b
  Symphytum officinale L. Symphytum officinale L.   26, 40, 48 Cave1963 Gadella & Klip. 1963
  Symphytum officinale L. Symphytum officinale L.   [48]U CromoCat 2015 Gadella, Th. W. J. & Kliphuis, E. (1972). Studies in chromosome numbers of Yugoslavian Angiosperms. Acta Bot. Croat. 31: 91-103.
  Symphytum officinale L. Symphytum officinale L.   24 CromoCat 2015 Gadella, T. W. J. & Kliphuis, E. (1978). Cytotaxonomic studies in the genus Symphytum. VlII. Chromosome numbers and classification of ten European species. Proc. K. Nederl. Akad. Wetensch. Amsterdam, Ser. C, 81: 162-172
  Symphytum officinale L. Symphytum officinale L.   24 CromoCat 2015 Huizing, H. J. (1985). Phytochemistry, systematics and biogenesis of pyrrolizidine alkaloids of Symphytum taxa. Thesis, Meppel.
  Symphytum officinale L. Symphytum officinale L.   24+0-4B CromoCat 2015 Bottega, S., Garbari, F. & Peruzzi, L. (2001). Mediterranean chromosome number reports-11. Fl. Medit. 11: 436-439.
  Symphytum officinale L. Symphytum officinale L.   40 CromoCat 2015 Huizing, H. J. (1985). Phytochemistry, systematics and biogenesis of pyrrolizidine alkaloids of Symphytum taxa. Thesis, Meppel.
  Symphytum officinale L. Symphytum officinale L.   40-48 CromoCat 2015 Gadella, Th. W. J. & Kliphuis, E. (1984). Population variability, hybridization and introgression in Symphytum officinale L. in the Netherlands Botanische Jahrbücher für Systematik, Pflanzengeschichte und Pflanzengeographie 104: 519-536
  Symphytum officinale L. Symphytum officinale L.   48 CromoCat 2015 Gadella, Th. W. J. & Kliphuis, E. (1972). Studies in chromosome numbers of Yugoslavian Angiosperms. Acta Bot. Croat. 31: 91-103.
  Symphytum officinale L. Symphytum officinale L.   48 CromoCat 2015 Gadella, Th. W. J., Kliphuis, E. & Kramer, K. U. (1970). Zytotaxonomische untersuchungen an blutenpflanzen aus dem Osten Osterreichs. Wiss. Arbeiten Burgenland 44: 187-195.
  Symphytum officinale L. Symphytum officinale L.   48 CromoCat 2015 Jaarsma, T. A., Lohmanns, E., Gadella, T. W. J. & Malingré, T. M. (1989). Chemotaxonomy of the Symphytum officinale agg. (Boraginaceae). Pl. Syst. Evol. 167(3/4): 113-127.
  Symphytum officinale L. Symphytum officinale L.   48 CromoCat 2015 Krahulcová, A. (1988). Selected chromosome counts of the Czechoslovak flora I. Folia Geobot. Phytotax. 23: 375-381.
  Symphytum officinale L. Symphytum officinale L.   48 CromoCat 2015 Loon van, J. C. & Kieft, B. (1980a). Reports [In Löve, A. (ed.), IOPB chromosome numbers reports LXVIII]. Taxon 29(4): 538-542. si
  Symphytum officinale L. Symphytum officinale L.   48 CromoCat 2015 Markova, M. & Ivanova, P. (1970). Karyologische Untersuchung der Vertreter der fam. Boraginaceae, Labiateae und Scrophulariaceae in Bulgarien. Bull. Inst. Bot. 20: 93-98.
  Symphytum officinale L. Symphytum officinale L.   48 CromoCat 2015 Peev, D. (1975b). Chromosome numbers and karyotypes. Compt. Rend. Acad. Bulg. Sci. 28: 521-523.
  Symphytum officinale L. Symphytum officinale L.   56 CromoCat 2015 Huizing, H. J. (1985). Phytochemistry, systematics and biogenesis of pyrrolizidine alkaloids of Symphytum taxa. Thesis, Meppel.
!   Symphytum officinale None Symphytum officinale L.   48 CromoCat 2015 Markova, M. & Ivanova, P. (1974). Reports [In Löve, A. (ed.), IOPB chromosome numbers reports XLIV]. Taxon 23(2/3): 378.
  Symphytum peregrinum Ledeb. Symphytum officinale L.   40 IPCN online Wang, G., Z. Li & Q. Meng. 1983. An observation of chromosome numbers of Comfrey from different sources. Acta Phytotax. Sin. 21: 55–59.
  Symphytum peregrinum Ledeb. Symphytum officinale L.   45, 65 IPCN online SHIRATO, K., T. Shintani, G. Nakanishi & A. Kamizy;afo. 1985. Chromosome numbers in genus Symphytum. Chromosome Inf. Serv. 38: 21–23.
  Symphytum peregrinum Ledeb. Symphytum officinale L.   40 IPCN online Huizing, H. J. 1985. Phytochemistry, systematics and biogenesis of pyrrolizidine alkaloids of Symphytum taxa. Thesis, Meppel. ý.
  Symphytum peregrinum Ledeb. Symphytum officinale L.   40 IPCN online Gadella, T. W., E. KLIPHUIS & H. J. Huizing. 1983. Cyto- and chemotaxonomical studies on the sections Officinalia and Coerulea of the genus Symphytum. Bot. Helv. 93: 169–192.
  Symphytum peregrinum Ledeb. Symphytum officinale L.   36 IPCN online Probatova. 2006. Chromosome numbers of plants of the Primorsky Territory, the Amur River basin and Magadan region. Bot. Žhurn. (Moscow & Leningrad) 91(3): 491–509.
  Symphytum peregrinum Ledeb. Symphytum officinale L.   40 IPCN72 GVINIASHVILI, TS. N. 1972. Some data on the karyology of Caucasic species of Symphytum L. with respect to their taxonomy. (In Russian). Bot. Zurn. 57: 1120-1126.
!   Symphytum peregrinum Ledeb. Symphytum officinale L.   36 Darlington, 1955 Maude 1939
  Symphytum peregrinum Ledeb. Symphytum officinale L.   40 IAPT/IOPB IAPT/IOPB Chromosome Data 19